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The single-player story follows Carl 'CJ' Johnson, who returns home to Los Santos after learning of his mother's murder. The state of San Andreas consists of three metropolitan cities: Los Santos, based on Los Angeles San Fierro, based on San Francisco and Las Venturas, based on Las Vegas. The game is set within the fictional US state of San Andreas, which is heavily based on California and Nevada.

CJ finds his old friends and family in disarray, and over the course of the game he attempts to re-establish his old gang, clashes with corrupt cops, and gradually unravels the truth behind his mother's murder.